 Barnes faces a race to be fit for the Test against England on 7 November
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Australia vice-captain Berrick Barnes will miss Saturday's Bledisloe Cup game against New Zealand in Tokyo after suffering an ankle injury in training.
X-rays showed Barnes has not broken the joint but it is not known if he can continue on Australia's tour, which heads to the British Isles after Japan.
In his absence, Australia move Adam Ashley-Cooper to inside centre with James O'Connor coming in at full-back.
"It's unfortunate but we are moving forward," said lock James Horwill.
"We're confident the guys in there are going to be more than able to do their jobs."
Australia have lost their last six meetings against their trans-Tasman rivals, including a 33-6 thrashing in September in Wellington.
With O'Connor's recall, Wallabies coach Robbie Deans makes five changes to his starting XV from that game, with wingers Digby Ioane and Peter Hynes, centre Ryan Cross and number eight Wycliff Palu the other new faces.
The All Blacks have made three changes, with winger Sitiveni Sivivatu, number eight Rodney So'oialo and centre Conrad Smith set to start in Tokyo.
Australia's Grand Slam tour of the British Isles gets under way with a Test against England on 7 November, followed by games against Ireland, Scotland and Wales on consecutive weekends.
Australia: Ashley-Cooper; Hynes, Cross, Barnes, Ioane; Giteau, Genia; Robinson, Polota-Nau, Alexander, Horwill, Chisholm, Elsom (capt), Pocock, Palu.
Replacements: Moore, Dunning, Mumm, Smith, Burgess, Mitchell, O'Connor
New Zealand: Muliaina; Jane, Smith, Nonu, Sivivatu; Carter, Cowan; Woodcock, Hore, Tialata, Thorn, Donnelly, Thomson, McCaw (capt), So'oialo.
Replacements: Corey Flynn, John Afoa, Jason Eaton, Kieran Read, Brendon Leonard, Stephen Donald, Tamati Ellison.
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